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However, it is precisely climate impacts in combination with human activity that are compromising the ability of ecosystems to provide these services

He has made a reputation for his work on vision and visual ecology in the marine environment, and is particularly interested in how vision has been matched to different marine habitats (e.g., shallow and brightly lit colorful reefs or the dim blue featureless world of the deep sea), and to the different visual needs of animals (e.g., in the context of mating, defense and predation)

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This is converted back to glutathione by the enzyme glutathione disulfide reductase by the molecule NADPH, i.e