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10.1016/j.abb.2012.04.014 Vido K, Diemer H, Van Dorsselaer A, Leize E, Juillard V, Gruss A, Gaudu P: Roles of thioredoxin reductase during the aerobic life of Lactococcus lactis

It is worth noting a point of ongoing scholarly discussion around this origin story: BPC-157, as studied, is a fully synthetic sequence, and its status as a naturally occurring fragment of a larger native protein has been questioned in later reviews

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Room temperature storage exposes peptides to atmospheric oxygen, and higher temperatures increase oxidation rates