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He received a VMD from the University of Pennsylvania in 1994 and completed a one year internship (small animal medicine and surgery) at The Animal Medical Center (NY, NY) in 1995, worked in small animal private practice in Maryland and Pennsylvania from 19952000, completed a three year residency in radiology from the University of Pennsylvania School of Veterinary Medicine in 2003, and completed a one year staff veterinarian position at the University of Pennsylvania in 2004

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Source of (an) (essential) amino acid(s) involved in muscle protein synthesis (IOM 2005)