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Glutamine has a versatile role in cell metabolism, participating in tricarboxylic acid (TCA) cycle supplementation and the biosynthesis of nucleotides, glutathione (GSH), and other nonessential amino acids

Glutamine is a neutral amino acid and thus does not induce a negative charge burden in the mitochondrial matrix, which is already more negatively charged than the cytosol

Importantly, nuclear PKM2 also exhibits protein kinase activity to phosphorylate histone H3 at T11, which is essential for the dissociation of HDAC3 from the c-Myc promoter to initiate c-Myc transcription (Yang et al., 2014)

Consequently, the fluorescence ratio between emission at 510 nm and excitation either at 488 nm or 405 nm can be used as an indicator of the relative amount of oxidized/reduced roGFP